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Table 2.

Comparison of parameters of behaviour between laps

A
BehaviourGroup1st lap2nd lap3rd lap4th lapGender effect (F1,10; P)Lap effect (F1,10; P)
Distance travelled in a lap (m) Males 21.2±5.7 15.6±3.1 15.2±4.4 9.6±1.2 37.8; <0.0001 1.9; 0.14 
 Females 49.8±13.1 27.8±5.4 29.4±9.5 29.6±8.9   
Lap duration (min) Males 4.4±1.4 1.9±0.5 1.9±0.8 1.2±0.4 8.7; 0.014 3.2; 0.036 
 Females 6.7±1.3 3.7±0.9 3.9±1.4 4.6±1.9   
Speed of travelling in a lap (cm s–1)2,3,4 Males 9.7±1.7 14.8±1.7 17.2±3.0 16.8±3.5 0.2; 0.697 5.5; 0.004 
 Females 12.1±1.6 14.5±2.2 14.7±2.2 13.1±2.1   
Inter-stop distance (m)3 Males 0.8±0.3 1.1±0.3 1.8±0.6 1.5±0.5 0.7; 0.225 3.7; 0.022 
 Females 0.7±0.1 0.9±0.2 1.0±0.2 0.9±0.2   
Number of stops in a lap3,4 Males 42.5±13.8 19.3±6.2 18.2±9.7 11.3±5.1 5.9; 0.035 4.4; 0.011 
 Females 77.0±25.6 37.7±8.6 35.5±12.3 41.2±15.2   
Backward walks (retreats) in a lap Males 12.8±5.2 2.5±1.1 4.0±2.7 1.8±1.1 2.2; 0.167 4.12; 0.0146 
 Females 12.3±4.9 8.0±3.5 7.3±3.5 8.0±3.0   
Reversals of progress direction (retraces) in a lap3 Males 3.0±0.9 0.8±0.6 1.0±0.7 0.7±0.4 29.5; 0.0003 4.8; 0.0078 
 Females 11.5±3.4 5.2±1.5 3.7±1.1 5.3±2.6   
Incidence of facing the centre in a lap3,4 Males 21.0±7.7 11.7±5.3 7.7±4.1 4.8±1.8 1.7; 0.219 6.5; 0.0016 
 Females 31.2±12.6 12.7±3.7 11.3±4.4 14.8±5.6   
       
Behaviour Group 1st visit 2nd visit 3rd visit 4th visit Gender effect (F1,10; PLap effect (F1,10; P
Speed within the corner zone (cm s–1Males 17.5±3.2 21.7±2.3 26.1±2.8 23.7±3.0 1.9; 0.199 6.9; 0.001 
 Females 20.1±2.6 27.4±2.3 28.1±3.7 29.7±3.9   
A
BehaviourGroup1st lap2nd lap3rd lap4th lapGender effect (F1,10; P)Lap effect (F1,10; P)
Distance travelled in a lap (m) Males 21.2±5.7 15.6±3.1 15.2±4.4 9.6±1.2 37.8; <0.0001 1.9; 0.14 
 Females 49.8±13.1 27.8±5.4 29.4±9.5 29.6±8.9   
Lap duration (min) Males 4.4±1.4 1.9±0.5 1.9±0.8 1.2±0.4 8.7; 0.014 3.2; 0.036 
 Females 6.7±1.3 3.7±0.9 3.9±1.4 4.6±1.9   
Speed of travelling in a lap (cm s–1)2,3,4 Males 9.7±1.7 14.8±1.7 17.2±3.0 16.8±3.5 0.2; 0.697 5.5; 0.004 
 Females 12.1±1.6 14.5±2.2 14.7±2.2 13.1±2.1   
Inter-stop distance (m)3 Males 0.8±0.3 1.1±0.3 1.8±0.6 1.5±0.5 0.7; 0.225 3.7; 0.022 
 Females 0.7±0.1 0.9±0.2 1.0±0.2 0.9±0.2   
Number of stops in a lap3,4 Males 42.5±13.8 19.3±6.2 18.2±9.7 11.3±5.1 5.9; 0.035 4.4; 0.011 
 Females 77.0±25.6 37.7±8.6 35.5±12.3 41.2±15.2   
Backward walks (retreats) in a lap Males 12.8±5.2 2.5±1.1 4.0±2.7 1.8±1.1 2.2; 0.167 4.12; 0.0146 
 Females 12.3±4.9 8.0±3.5 7.3±3.5 8.0±3.0   
Reversals of progress direction (retraces) in a lap3 Males 3.0±0.9 0.8±0.6 1.0±0.7 0.7±0.4 29.5; 0.0003 4.8; 0.0078 
 Females 11.5±3.4 5.2±1.5 3.7±1.1 5.3±2.6   
Incidence of facing the centre in a lap3,4 Males 21.0±7.7 11.7±5.3 7.7±4.1 4.8±1.8 1.7; 0.219 6.5; 0.0016 
 Females 31.2±12.6 12.7±3.7 11.3±4.4 14.8±5.6   
       
Behaviour Group 1st visit 2nd visit 3rd visit 4th visit Gender effect (F1,10; PLap effect (F1,10; P
Speed within the corner zone (cm s–1Males 17.5±3.2 21.7±2.3 26.1±2.8 23.7±3.0 1.9; 0.199 6.9; 0.001 
 Females 20.1±2.6 27.4±2.3 28.1±3.7 29.7±3.9   

Significant differences are shown in bold

None of the interactions (lap × gender) were significant (data not shown)

There was no significant difference between laps 2, 3 and 4 (Tukey HSD test). Superscript numbers in the `Behaviour' column indicate the number of the lap from which the first lap significantly differed (Tukey HSD test)

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